welke zijn de vaste artributen of vaste menages die je op tafel zet.

zout , peper -vaas- met bloemen-kaars

Materialen die gebruikt worden in restaurant

Hinter den Bergen Zwei Bilder sagen mehr als tausend Worte Wolfgang Ullrich Wer über Erfahrung mit Bildern verfügt, weiss, wie unerwartet und wie stark sie aufeinander reagieren können. So uninteressant ein Bild für sich alleine sein mag, so komisch, zynisch, erhellend oder erhebend wird es oft, wenn man es mit anderen Bildern kombiniert. Ähnliches kennt man sonst am ehesten vom Chemieunterricht: Zuerst harmlose Stoffe entfalten ungeahnte Energien und Effekte, sobald sie zusammengebracht werden. Besonders verblüffend ist immer wieder, was alles durch die Verbindung von nur zwei Bildern passieren kann. Einmal werden sie zum Gegensatzpaar, in einem anderen Fall bestätigen und verstärken sie sich gegenseitig – oder sie variieren ein Thema, erzählen auf knappste Weise eine Geschichte, geben in ihrer Verschiedenheit ein Rätsel auf. In der Geschichte der Kunst, aber ebenso in der Propaganda oder Werbung nutzte man die Aussagekraft von Bildpaaren häufig: vom Diptychon in der christlichen Tradition des Andachtsbilds bis hin zu plakativen Vorher-nachher-Bildern, mit denen ein Haarwuchsmittel angepriesen oder ein kultureller Niedergang beklagt wird. In der Moderne, in der Bilder allgemein verfügbar geworden sind, ist das Kombinieren von Bildern sogar mindestens so wichtig geworden wie das Machen neuer Bilder. Die wohl subtilste Form des Bildpaars taucht aber vor allem in früheren Jahrhunderten auf. Es handelt sich dabei um Pendants. Sie ergeben sich daraus, dass man zwei Bilder zusammenfügt, die ein verwandtes Sujet zeigen, es aber unterschiedlich behandeln, also aus jeweils anderer Perspektive, in verschiedenen Stimmungen oder mit wechselnden Konnotationen darstellen. Die Blütezeit der Pendants war das 18. Jahrhundert, sie wurden aber bereits von Claude Lorrain und auch noch von Caspar David Friedrich virtuos eingesetzt.[1] Viele Pendants stellten aber nicht einmal die Künstler selbst zusammen; vielmehr wurden sie in den Werkstätten und Verkaufsräumen von Druckgraphikern arrangiert. Diese erkannten nämlich, dass sich Kupferstiche oft besser paarweise als alleine verkaufen ließen, weil sie dem Publikum dann mehr Stoff zum Räsonieren boten. So wurde es sogar üblich, Stiche von Werken verschiedener Künstler zu kombinieren, und selbst Bilder, die zu unterschiedlichen Zeiten entstanden waren, ließen sich als Pendants vereint wiederfinden.[2] Der Tiroler Fotograf Lois Hechenblaikner hat diese Praxis nicht nur neu aufgegriffen, sondern sie um eine zusätzliche – entscheidende – Dimension bereichert. In Gegenüberstellungen, an denen er seit dem Jahr 2000 arbeitet, präsentiert er jeweils ein Bild in Schwarz-Weiß – meist ein Foto aus dem reichen Nachlass des Agraringenieurs Armin Kniely (1907-1998) – neben einem Farbbild, das er selbst fotografiert hat. Die unterschiedlichen Entstehungszeiten der Bilder sind damit, anders als bei den Pendants des 18. Jahrhunderts, bereits auf den ersten Blick erkennbar und auch Thema: Lois Hechenblaikner geht es darum, sichtbar zu machen, wie sich eine Region – fast alle Motive stammen aus Tirol – innerhalb von nur zwei Generationen verändert hat. Wo man in den 1930er und selbst noch in den 1960er Jahren Bergbauern bei ihrer mühsamen Feldarbeit aufnehmen sowie ein ritualisiertes Brauchtum dokumentierten konnte, sind mittlerweile der Tourismus und die Formate der Eventkultur am prägendsten geworden. Doch statt nur einen Strukturwandel festzuhalten, besteht Hechenblaikners besondere Fähigkeit darin, in jeder Gegenüberstellung zwei formal analoge Szenerien aufeinander zu beziehen. Erst so werden aus seinen Bildpaaren auch Pendants – zwei Fotos, die trotz aller Unterschiede jeweils genügend Gemeinsames haben, um zum Vergleich herauszufordern. Die Bildpaare gehen bei Hechenblaikner auch immer über das bloß Assoziative hinaus, das in der Kunst sonst beliebt ist, weil das Arrangement dann eine Vieldeutigkeit verheißt und verrätselt wirkt (oft aber nur beliebig ist). Vielmehr empfindet man seine Pendants als geradezu unheimlich – als unglaublich und grotesk –, weil die von ihm entdeckten Ähnlichkeiten so stark sind. Daher überlegt man auch, ob es sich bei den Farbfotos nicht vielleicht um direkte Remakes der Schwarz-weiß-Motive handelt. Doch ist schnell zu erkennen, dass Lois Hechenblaikner seine Bilder nicht eigens inszeniert: Da fast immer Menschen, nicht selten sogar größere Gruppen darauf zu sehen sind, wäre es zu aufwändig, für einen einzelnen Fotografen gar unmöglich, die Szenen zu stellen. Vielmehr ist es Hechenblaikners hervorragendem Bildgedächtnis zu verdanken, wenn er bei seinen Streifzügen durch das Land immer wieder Situationen entdeckt, die denen auf alten Fotos genau entsprechen: Wie einst ein Bauer Jauche auf die Felder spritzte, so schießt man jetzt Schnee aus Kanonen, die Stecken, um die man früher das Heu zum Trocknen wickelte, ähneln verblüffend Handy-Masten, und wo die Landwirte einst stolz ihre Schafe präsentierten, posieren heute Golfspieler mit ihren Trolleys. Die Botschaft der Pendants ist somit eine doppelte: Sie künden sowohl von den gewaltigen Umbrüchen in den Alpen während der letzten Jahrzehnte als auch davon, dass alle Veränderungen doch nur scheinbar sind, ja dass sich lediglich die Hüllen und Akzidenzien gewandelt haben, in denen dieselben Muster geradezu zeitlos zur Geltung kommen. Man könnte darin eine Bestätigung für das sehen, was Aby Warburg als Pathosformeln bezeichnet hat, nämlich Gesten und Konstellationen, die über Epochen und sogar über Kulturen hinweg immer wieder auftauchen und die damit den Status anthropologischer Konstanten besitzen. Wollte Warburg diese Formeln mithilfe eines Atlasses erforschen, der Bilder unterschiedlichster Herkunft mit jeweils verwandten Sujets auf Tableaus versammelte[3], so läßt sich das vorliegende Buch von Lois Hechenblaikner als Tiroler Variante eines ähnlichen Projekts würdigen. Doch während Warburg, getrieben von metaphysischem Interesse und auf der Suche nach so etwas wie Archetypen menschlicher Existenz, auf den einzelnen Tafeln seines Atlasses weit ausholt und auch Bilder zusammenbringt, die zumindest auf den ersten Blick nur wenig gemeinsam haben, ist Hechenblaikners Arbeit ungleich konzentrierter. Da er immer nur Bildpaare zeigt, denen zudem allen dasselbe Schema zugrunde liegt, wird der Betrachter viel stärker geführt. Jede Doppelseite gerät bei ihm zu einer neuen Pointe. Doch verhindert die Form des Pendants, dass mit dem Erfassen der Pointe schon alles vorbei ist. Im Gegenteil fordert sie zur weiteren Reflexion heraus, und das Spiel von Identität und Differenz, das Lois Hechenblaikner mit jeder Gegenüberstellung auf die Spitze treibt, eröffnet dafür den Raum. Viel mehr als einzelne Bilder, mehr aber auch als andere Formen von Bildpaaren verlangen Pendants sogar ausdrücklich aktive Rezipienten, die diesen Raum mit ihren Gedanken füllen. Sie können dann entweder darüber sinnieren, wieso sich zwei vermeintlich so unterschiedliche Lebenswelten wie Landwirtschaft und Tourismus, Religion und Sport, Brauchtum und Kommerz in so ähnlichen Bildformeln ausdrücken, oder sie können der Überlegung nachgehen, wie sich eine Region innerhalb kurzer Zeit so stark verändern konnte, dass alle Tätigkeiten und Situationen komplett durch andere Tätigkeiten und Situationen ersetzt wurden. Ist also einmal die Ähnlichkeit des Unterschiedlichen das Faszinosum, so das andere Mal die Andersheit des Gleichen. Jeweils aber muss der Rezipient beides – Identität und Differenz – zusammendenken. Er kann sich dabei auf ein einzelnes Beispiel konzentrieren und Überlegungen zur jeweiligen Rolle etwa von Schafen und Trolleys – zu ihrer Eignung als Symbole für eine Lebensform – anstellen. Er kann sich aber auch von der Summe der Bildpaare anregen lassen und so zu Theoriebildung animiert werden, um für das ihm Dargebotene eine Erklärung zu finden. In einem Text spricht Lois Hechenblaikner selbst davon, durch die Bildpaare solle „im Kopf des Betrachters ein so starkes drittes Bild entstehen, dass ihm förmlich ‚das Licht aufgeht’“. Doch sind es – je nach Interesse, Haltung und Vorbildung – durchaus unterschiedliche Lichter, die aufgehen können. Während Hechenblaikner die Gegenüberstellungen zuerst mit dem Ziel anlegte, die Veränderung Tirols und des Alpenraumes als Verlustgeschichte zu erzählen, ist es mit ihnen genauso möglich, die verschwundene Welt nachträglich zu dekonstruieren. Läßt der Wechsel vom Schwarz-weiß zur Farbe einerseits den Schluss zu, dass sich in der Gegenwart eine brutale Marktlogik durchgesetzt hat, derzufolge alles mit möglichst starken Reizen – mit Grellheit und Obszönität – auf sich aufmerksam macht, so kann man daraus andererseits also auch die Mahnung ableiten, auf das Vergangene nicht nur deshalb sentimental zu blicken, weil es durch die einheitliche Tönung verfremdet erscheint. Obwohl das Leben der Bergbauern, ihre familiäre Bindung, ihr Gottvertrauen und ihre dem Rhythmus der Jahreszeiten gehorchende Arbeit idyllisch wirken mag, könnte das alles vielmehr ähnlich – genauso? – derb, banal und kalt gewesen sein wie die heutige von Unterhaltungsindustrie, Animiergewerbe und Konsum bestimmte Welt des Massentourismus. Oder muss es sich doch eindeutig um einen Verfall handeln, wenn allenthalben natürliche Materialien durch Kunststoffe ausgetauscht, freie Blicke durch Werbetafeln verstellt, beschauliche Szenerien von Massenevents abgelöst, Ernst und Strenge durch Albernheit und Ausgelassenheit ersetzt wurden? Aber ist es nicht auch ein Zeichen von Freiheit und Wohlstand – und damit sogar ein Fortschritt –, wenn die Menschen ihre Launen heute ausleben können und nicht länger ein enges, allein von Armut und Notwendigkeiten bestimmtes Leben führen müssen? Selbst wenn man klar für eine Deutung optiert und sich entweder als Kulturpessimist oder als Fürsprecher einer grundsätzlichen Gleichwertigkeit verschiedener Epochen oder aber als Fortschrittseuphoriker bekennt, kann man die jeweils alternativen Interpretationen wohl nie ganz ausblenden. Je besser ein Bildpaar ist, desto stärker sind vielmehr zugleich die jeweils anderen Sichtweisen präsent. Es gehört sogar geradezu zum Wesen von Pendants (wie ihr Name, abgeleitet vom lateinischen ‚pendere‘, bereits verrät), zu einem Abwägen zu verleiten und feste Positionen in die Schwebe zu bringen. Pendants sind also die ideale Bildform für Skeptiker, die sich ungern festlegen und immer gerne mindestens eine zweite Sichtweise parat haben. Zwei Bilder, so wie Lois Hechenblaikner sie aufeinander bezieht, sagen daher tatsächlich mehr als tausend Worte, mit denen nämlich jeweils nur eine einzige Position formuliert werden kann: Satz und Gegensatz in sich vereinend, verführen Pendants zum philosophischen Denken – zu einem nie abschließbaren Nachdenken über Geschichte, Menschen, Kultur. Das Schwanken zum Prinzip erhebend, sind sie die besten Stimulanzien des Geistes.

Figure 18-11 represents the amount of energy stored as organic material in each trophic level in an ecosystem. The pyramid shape of the diagram indicates the low percentage of energy transfer from one level to the next. On average, 10 percent of the total energy consumed in one trophic level is incor- porated into the organisms in the next. Why is the percentage of energy transfer so low? One reason is that some of the organisms in a trophic level escape being eaten. They eventually die and become food for decomposers, but the energy contained in their bodies does not pass to a higher trophic level. Even when an organism is eaten, some of the molecules in its body will be in a form that the consumer cannot break down and use. For example, a cougar cannot extract energy from the antlers, hooves, and hair of a deer. Also, the energy used by prey for cellu- lar respiration cannot be used by predators to synthesize new bio- mass. Finally, no transformation or transfer of energy is 100 percent efficient. Every time energy is transformed, such as during the reactions of metabolism, some energy is lost as heat. Limitations of Trophic Levels The low rate of energy transfer between trophic levels explains why ecosystems rarely contain more than a few trophic levels. Because only about 10 percent of the energy available at one trophic level is transferred to the next trophic level, there is not enough energy in the top trophic level to support more levels. Organisms at the lowest trophic level are usually much more abundant than organisms at the highest level. In Africa, for exam- ple, you will see about 1,000 zebras, gazelles, and other herbivores for every lion or leopard you see, and there are far more grasses and shrubs than there are herbivores. Higher trophic levels con- tain less energy, so, they can support fewer individuals.A population is a group of organisms that belong to the same species and live in a particular place at the same time. All of the bass living in a pond during a certain period of time make up a pop- ulation because they are isolated in the pond and do not interact with bass living in other ponds. The boundaries of a population may be imposed by a feature of the environment, such as a lake shore, or they can be arbitrarily chosen to simplify a study of the population. The humans shown in Figure 19-1 are part of the pop- ulation of a city. The properties of populations differ from those of individuals. An individual may be born, it may reproduce, or it may die. A population study focuses on a population as a whole—how many individuals are born, how many die, and so on. Population Size A population’s size is the number of individuals that the population contains. Size is a fundamental and important population property but can be difficult to measure directly. If a population is small and composed of immobile organisms, such as plants, its size can be determined simply by counting individuals. Often, though, individ- uals are too abundant, too widespread, or too mobile to be counted easily, and scientists must estimate the number of individuals in the population. Suppose that a scientist wants to know how many oak trees live in a 10 km2 patch of forest. Instead of searching the entire patch of forest and counting all the oak trees, the scientist could count the trees in a smaller section of the forest, such as a 1 km2 area. The scientist could then use this value to estimate the population of the larger area. SECTION 1 OBJECTIVES ● Describe the main properties that scientists measure when they study populations. ● Compare the three general patterns of population dispersion. ● Identify the measurements used to describe changing populations. ● Compare the three general types of survivorship curves. VOCABULARY population population density dispersion birth rate death rate life expectancy age structure survivorship curve FIGURE 19-1 A population can be widely distributed, as Earth’s human population is, or confined to a small area, as species of fish in a lake are. Copyright © by Holt, Rinehart and Winston. All rights reserved. 382 CHAPTER 19 If the small patch contains 25 oaks, an area 10 times larger would likely contain 10 times as many oak trees. A similar kind of sampling technique might be used to estimate the size of the pop- ulation shown in Figure 19-2. To use this kind of estimate, the sci- entist must assume that the distribution of individuals in the entire population is the same as that in the sampled group. Estimates of population size are based on many such assumptions, so all esti- mates have the potential for error. Population Density Population density measures how crowded a population is. This measurement is always expressed as the number of individuals per unit of area or volume. For example, the population density of humans in the United States is about 30 people per square kilome- ter. Table 19-1 shows the population sizes and densities of humans in several countries in 2003. These estimates are calculated for the total land area. Some areas of a country may be sparsely popu- lated, while other areas are very densely populated. Dispersion A third population property is dispersion (di-SPUHR-zhuhn). Dispersion is the spatial distribution of individuals within the popu- lation. In a clumped distribution, individuals are clustered together. In a uniform distribution, individuals are separated by a fairly con- sistent distance. In a random distribution, each individual’s location is independent of the locations of other individuals in the popula- tion. Figure 19-3 illustrates the three possible patterns of dispersion. Clumped distributions often occur when resources such as food or living space are clumped. Clumped distributions may also occur because of a species’ social behavior, such as when animals gather into herds or flocks. Uniform distributions may result from social behavior in which individuals within the same habitat stay as far away from each other as possible. For example, a bird may locate its nest so as to maximize the distance from the nests of other birds. These migrating wildebeests in East Africa are too numerous and mobile to be counted. Scientists must use sampling methods at several locations to monitor changes in the population size of the animals. FIGURE 19-2 TABLE 19-1 Population Size and Density of Some Countries Population size Population density Country (in millions) (in individuals/km2) China 1,289 135 India 1,069 325 United States 292 30 Russia 146 8 Japan 128 337 Mexico 105 54 Kenya 32 54 Australia 20 3 dispersion from the Latin dis-, meaning “out,” and spargere, meaning “to scatter” Word Roots and Origins Copyright © by Holt, Rinehart and Winston. All rights reserved. POPULATIONS 383 The social interactions of birds called gannets, which are shown in Figure 19-3b, result in a uniform distribution. Each gannet chooses a small nesting area on the coast and defends it from other gannets. In this way, each gannet tries to maximize its distance from all of its neighbors, which causes a uniform distribution of individuals. Few populations are truly randomly dispersed. Rather, they show degrees of clumping or uniformity. The dispersion pattern of a population sometimes depends on the scale at which the popu- lation is observed. The gannets shown in Figure 19-3b are uni- formly distributed on a scale of a few meters. However, if the entire island on which the gannets live is observed, the distribution appears clumped because the birds live only near the shore. POPULATION DYNAMICS All populations are dynamic—they change in size and composition over time. To understand these changes, scientists must know more than the population’s size, density, and dispersion. One important measure is the birth rate, the number of births occur- ring in a period of time. In the United States, for example, there are about 4 million births per year. A second important measure is the death rate, or mortality rate, which is the number of deaths in a

CARBOHYDRATES Carbohydrates are organic compounds composed of carbon, hydrogen, and oxygen in a ratio of about one carbon atom to two hydrogen atoms to one oxygen atom. The number of carbon atoms in a carbohydrate varies. Some carbohydrates serve as a source of energy. Other carbohydrates are used as structural materials. Carbohydrates can exist as monosaccharides, disaccharides, or polysaccharides. Monosaccharides A monomer of a carbohydrate is called a monosaccharide (MAHN-oh-SAK-uh-RIED). A monosaccharide—or simple sugar— contains carbon, hydrogen, and oxygen in a ratio of 1:2:1. The gen- eral formula for a monosaccharide is written as (CH2O)n, where n is any whole number from 3 to 8. For example, a six-carbon mono- saccharide, (CH2O)6, would have the formula C6H12O6. The most common monosaccharides are glucose, fructose, and galactose, as shown in Figure 3-6. Glucose is a main source of energy for cells. Fructose is found in fruits and is the sweetest of the monosaccharides. Galactose is found in milk. Notice in Figure 3-6 that glucose, fructose, and galactose have the same molecular formula, C6H12O6, but differing structures. The different structures determine the slightly different properties of the three compounds. Compounds like these sugars, with a single chemical formula but different structural forms, are called isomers (IE-soh-muhrz). SECTION 2 OBJECTIVES ● Distinguish between monosaccharides, disaccharides, and polysaccharides. ● Explain the relationship between amino acids and protein structure. ● Describe the induced fit model of enzyme action. ● Compare the structure and function of each of the different types of lipids. ● Compare the nucleic acids DNA and RNA. VOCABULARY carbohydrate monosaccharide disaccharide polysaccharide protein amino acid peptide bond polypeptide enzyme substrate active site lipid fatty acid phospholipid wax steroid nucleic acid deoxyribonucleic acid (DNA) ribonucleic acid (RNA) nucleotide C HO H C H OH C OH H C CH2OH H C H OH O Glucose C OH C O H OH C OH H CH2OH C H CH2OH Fructose C H HO C OH H C OH H C CH2OH H C H OH O Galactose Glucose, fructose, and galactose have the same chemical formula, but their structural differences result in different properties among the three compounds. FIGURE 3-6 Copyright © by Holt, Rinehart and Winston. All rights reserved. 56 CHAPTER 3 Disaccharides and Polysaccharides In living things, two monosaccharides can combine in a condensa- tion reaction to form a double sugar, or disaccharide (die-SAK-e-RIED). For example in Figure 3-4, the monosaccharides fructose and glu- cose can combine to form the disaccharide sucrose. A polysaccharide is a complex molecule composed of three or more monosaccharides. Animals store glucose in the form of the polysaccharide glycogen. Glycogen consists of hundreds of glucose molecules strung together in a highly branched chain. Much of the glucose that comes from food is ultimately stored in your liver and muscles as glycogen and is ready to be used for quick energy. Plants store glucose molecules in the form of the polysaccha- ride starch. Starch molecules have two basic forms—highly branched chains that are similar to glycogen and long, coiled, unbranched chains. Plants also make a large polysaccharide called cellulose. Cellulose, which gives strength and rigidity to plant cells, makes up about 50 percent of wood. In a single cellu- lose molecule, thousands of glucose monomers are linked in long, straight chains. These chains tend to form hydrogen bonds with each other. The resulting structure is strong and can be broken down by hydrolysis only under certain conditions. PROTEINS Proteins are organic compounds composed mainly of carbon, hydrogen, oxygen, and nitrogen. Like most of the other biological macromolecules, proteins are formed from the linkage of monomers called amino acids. Hair and horns, as shown in Figure 3-7a, are made mostly of proteins, as are skin, muscles and many biological catalysts (enzymes). Amino Acids There are 20 different amino acids, and all share a basic structure. As Figure 3-7b shows, each amino acid contains a central carbon atom covalently bonded to four other atoms or functional groups. A single hydrogen atom, highlighted in blue in the illustration, bonds at one site. A carboxyl group, —COOH, highlighted in green, bonds at a second site. An amino group, —NH2, highlighted in yel- low, bonds at a third site. A side chain called the R group, high- lighted in red, bonds at the fourth site. The main difference among the different amino acids is in their R groups. The R group can be complex or it can be simple, such as the CH3 group shown in the amino acid alanine in Figure 3-7b. The differences among the amino acid R groups gives different proteins very different shapes. The different shapes allow pro- teins to carry out many different activities in living things. Amino acids are commonly shown in a simplified way such as balls, as shown in Figure 3-7c. (a) Many structures, such as hair and horns are made of proteins. (b) Proteins are made up of amino acids. Amino acids differ only in the type of R group (shown in red) they carry. Polar R groups can dissolve in water, but nonpolar R groups cannot. (c) Amino acids have complex structures, so, in this and other textbooks, they are often simplified into balls. FIGURE 3-7 (b) Alanine (an amino acid) (c) Simplified version of amino acid CH3 H N OH C C H O H (a) Copyright © by Holt, Rinehart and Winston. All rights reserved. BIOCHEMISTRY 57 H H N C C OH H O H CH3 H2O Glycine Alanine H N OH C C H O H H H N C C H O H CH3 N OH C C H O H (a) (b) (a) The peptide bond (shaded blue) that binds amino acids together to form a polypeptide results from a condensation reaction that produces water. (b) Poly- peptides are commonly shown as a string of balls in this textbook and elsewhere. Each ball represents an amino acid. FIGURE 3-8 Substrate Products Enzyme 1 2 3 In the induced fit model of enzyme action, the enzyme can attach only to a substrate (reactant) with a specific shape. The enzyme then changes and reduces the activation energy of the reaction so reactants can become products. The enzyme is unchanged and is available to be used again. 3 2 1 FIGURE 3-9 Dipeptides and Polypeptides Figure 3-8a shows how two amino acids bond to form a dipeptide (die-PEP-TIED). In this condensation reaction, the two amino acids form a covalent bond, called a peptide bond (shaded in blue in Figure 3-8a) and release a water molecule. Amino acids often form very long chains called polypeptides (PAHL-i-PEP-TIEDZ). Proteins are composed of one or more polypep- tides. Some proteins are very large molecules, containing hun- dreds of amino acids. Often, these long proteins are bent and folded upon themselves as a result of interactions—such as hydrogen bonding—between individual amino acids. Protein shape can also be influenced by conditions such as temperature and the type of solvent in which a protein is dissolved. For exam- ple, cooking an egg changes the shape of proteins in the egg white. The firm, opaque result is very different from the initial clear, runny material. Enzymes Enzymes—RNA or protein molecules that act as biological catalysts—are essential for the functioning of any cell. Many enzymes are proteins. Figure 3-9 shows an induced fit model of enzyme action. Enzyme reactions depend on a physical fit between the enzyme molecule and its specific substrate, the reactant being catalyzed. Notice that the enzyme has folds, or an active site, with a shape that allows the substrate to fit into the active site. An enzyme acts only on a specific substrate because only that substrate fits into its active site. The linkage of the enzyme and substrate causes a slight change in the enzyme’s shape. The change in the enzyme’s shape weakens some chemical bonds in the substrate, which is one way that enzymes reduce activation energy, the energy needed to start the reaction. After the reaction, the enzyme releases the products. Like any catalyst, the enzyme itself is unchanged, so it can be used many times. An enzyme may not work if its environment is changed. For example, change in temperature or pH can cause a change in the shape of the enzyme or the substrate. If such a change happens, the reaction that the enzyme would have catalyzed cannot occur.

The endoplasmic reticulum (EN-doh-PLAZ-mik ri-TIK-yuh-luhm), abbre- viated ER, is a system of membranous tubes and sacs, called cisternae (sis-TUHR-nee). The ER functions primarily as an intracellu- lar highway, a path along which molecules move from one part of the cell to another. The amount of ER inside a cell fluctuates, depending on the cell’s activity. There are two types of ER: rough and smooth. The two types of ER are thought to be continuous. Rough Endoplasmic Reticulum The rough endoplasmic reticulum is a system of interconnected, flattened sacs covered with ribosomes, as shown in Figure 4-15. The rough ER produces phospholipids and proteins. Certain types of proteins are made on the rough ER’s ribosomes. These proteins are later exported from the cell or inserted into one of the cell’s own membranes. For example, ribosomes on the rough ER make digestive enzymes, which accumulate inside the endoplasmic retic- ulum. Little sacs or vesicles then pinch off from the ends of the rough ER and store the digestive enzymes until they are released from the cell. Rough ER is most abundant in cells that produce large amounts of protein for export, such as cells in digestive glands and antibody-producing cells. Smooth Endoplasmic Reticulum The smooth ER lacks ribosomes and thus has a smooth appear- ance. Most cells contain very little smooth ER. Smooth ER builds lipids such as cholesterol. In the ovaries and testes, smooth ER produces the steroid hormones estrogen and testosterone. In skeletal and heart muscle cells, smooth ER releases calcium, which stimulates contraction. Smooth ER is also abundant in liver and kidney cells, where it helps detoxify drugs and poisons. Long-term abuse of alcohol and other drugs causes these cells to produce more smooth ER. Increased amounts of smooth ER in liver cells is one of the factors that can lead to drug tolerance. As Figure 4-15 shows, rough ER and smooth ER form an interconnected network. Copyright © by Holt, Rinehart and Winston. All rights reserved. reticulum from the Latin rete, meaning “net”; reticulum means “little net” Word Roots and Origins The endoplasmic reticulum (ER) serves as a site of synthesis for proteins, lipids, and other materials. The dark lines in the photo represent the membranes of the ER, and the narrow lighter areas between the dark lines show the channels and spaces (cisternae) inside the ER. FIGURE 4-15 Smooth ER Ribosomes Rough ER Cisternae 82 CHAPTER 4 GOLGI APPARATUS The Golgi apparatus, shown in Figure 4-16, is another system of flattened, membranous sacs. The sacs nearest the nucleus receive vesicles from the ER containing newly made proteins or lipids. Vesicles travel from one part of the Golgi apparatus to the next and transport substances as they go. The stacked membranes modify the vesicle contents as they move along. The proteins get “address labels” that direct them to various other parts of the cell. During this modification, the Golgi apparatus can add carbohydrate labels to proteins or alter new lipids in various ways. VESICLES Cells contain several types of vesicles, which perform various roles. Vesicles are small, spherically shaped sacs that are surrounded by a single membrane and that are classified by their contents. Vesicles often migrate to and merge with the plasma membrane. As they do, they release their contents to the outside of the cell. Lysosomes Lysosomes (LIE-suh-SOHMZ) are vesicles that bud from the Golgi appa- ratus and that contain digestive enzymes. These enzymes can break down large molecules, such as proteins, nucleic acids, car- bohydrates, and phospholipids. In the liver, lysosomes break down glycogen in order to release glucose into the bloodstream. Certain white blood cells use lysosomes to break down bacteria. Within a cell, lysosomes digest worn-out organelles in a process called autophagy (aw-TAHF-uh-jee). Lysosomes are also responsible for breaking down cells when it is time for the cells to die. The digestion of damaged or extra cells by the enzymes of their own lysosomes is called autolysis (aw-TAHL-uh-sis). Lysosomes play a very important role in maintaining an organism’s health by destroying cells that are no longer functioning properly. Copyright © by Holt, Rinehart and Winston. All rights reserved. The Golgi apparatus modifies many cellular products and prepares them for export. FIGURE 4-16 CELL STRUCTURE AND FUNCTION 83 Peroxisomes Peroxisomes are similar to lysosomes but contain different enzymes and are not produced by the Golgi apparatus. Peroxisomes are abundant in liver and kidney cells, where they neutralize free radicals (oxygen ions that can damage cells) and detoxify alcohol and other drugs. Peroxisomes are named for the hydrogen peroxide, H2O2, they produce when breaking down alco- hol and killing bacteria. Peroxisomes also break down fatty acids, which the mitochondria can then use as an energy source. Other Vesicles Specialized peroxisomes, called glyoxysomes, can be found in the seeds of some plants. They break down stored fats to provide energy for the developing plant embryo. Some cells engulf material by surrounding it with plasma membrane. The resulting pocket buds off to become a vesicle inside the cell. This vesicle is called an endosome. Lysosomes fuse with endosomes and digest the engulfed material. Food vacuoles are vesicles that store nutrients for a cell. Contractile vacuoles are vesicles that can contract and dispose of excess water inside a cell. Protein Synthesis One of the major functions of a cell is the production of protein. The path some proteins take from synthesis to export can be seen in Figure 4-17. In step , proteins are assembled by ribosomes on the rough ER. Then, in step , vesicles transport proteins to the Golgi apparatus. In step , the Golgi modifies proteins and pack- ages them in new vesicles. In step , vesicles release proteins that have destinations outside the cell. In step , vesicles containing enzymes remain inside the cell as lysosomes, peroxisomes, endo- somes, or other types of vesicles. 5 4 3 2 1 Copyright © by Holt, Rinehart and Winston. All rights reserved. Proteins are assembled by ribosomes on the rough ER. Vesicles carry proteins from the rough ER to the Golgi apparatus. Proteins are modified in the Golgi apparatus and enter new vesicles. Some vesicles release their proteins outside the cell. Other vesicles remain in the cell and become lysosomes and other vesicles. Nucleus

CHARACTERISTICS OF LIFE The world is filled with familiar objects, such as tables, rocks, plants, pets, and automobiles. Which of these objects are living or were once living? What are the criteria for assigning something to the living world or the nonliving world? Biologists have established that living things share seven characteristics of life. These characteristics are organization and the presence of one or more cells, response to a stimulus (plural, stimuli), homeostasis, metabolism, growth and development, reproduction, and change through time. Organization and Cells Organization is the high degree of order within an organism’s internal and external parts and in its interactions with the living world. For example, compare an owl to a rock. The rock has a spe- cific shape, but that shape is usually irregular. Furthermore, differ- ent rocks, even rocks of the same type, are likely to have different shapes and sizes. In contrast, the owl is an amazingly organized individual, as shown in Figure 1-2. Owls of the same species have the same body parts arranged in nearly the same way and interact with the environment in the same way. Copyright © by Holt, Rinehart and Winston. All rights reserved. ORGANISM (Barn Owl) ORGAN (Owl’s Ear) TISSUE (Nervous Tissue Within the Ear) CELL (Nerve Cell) Every living organism has a level of organization. The different levels of organization for a complex multicellular organism, such as an owl, are shown in the figure below. FIGURE 1-2 THE SCIENCE OF LIFE 7 All living organisms, whether made up of one cell or many cells, have some degree of organization. A cell is the smallest unit that can perform all life’s processes. Some organisms, such as bacteria, are made up of one cell and are called unicellular (YOON-uh-SEL-yoo-luhr) organisms. Other organisms, such as humans or trees, are made up of multiple cells and are called multicellular (MUHL-ti-SEL-yoo-luhr) organisms. Complex multicellular organisms have the level of orga- nization shown in Figure 1-2. In the highest level, the organism is made up of organ systems, or groups of specialized parts that carry out a certain function in the organism. For example, an owl’s ner- vous system is made up of a brain, sense organs, nerve cells, and other parts that sense and respond to the owl’s surroundings. Organ systems are made up of organs. Organs are structures that carry out specialized jobs within an organ system. An owl’s ear is an organ that allows the owl to hear. All organs are made up of tissues. Tissues are groups of cells that have similar abilities and that allow the organ to function. For example, nervous tissue in the ear allows the ear to detect sound. Tissues are made up of cells. A cell must be covered by a membrane, contain all genetic information necessary for replication, and be able to carry out all cell functions. Within each cell are organelles. Organelles are tiny structures that carry out functions necessary for the cell to stay alive. Organelles contain biological molecules, the chemical compounds that provide physical structure and that bring about movement, energy use, and other cellular functions. All biological molecules are made up of atoms. Atoms are the simplest particle of an ele- ment that retains all the properties of a certain element. Response to Stimuli Another characteristic of life is that an organism can respond to a stimulus—a physical or chemical change in the internal or external environment. For example, an owl dilates its pupils to keep the level of light entering the eye constant. Organisms must be able to respond and react to changes in their environment to stay alive. ORGANELLE (Mitochondrion) BIOLOGICAL MOLECULE (Phospholipid) ATOM (Oxygen) cell from the Latin, cella meaning “small room,” or “hut” Word Roots and Origins www.scilinks.org Topic: Characteristics of Life Keyword: HM60257 mb06se_bios01.qxd 5/18/07 10:37 AM Page 7 8 CHAPTER 1 Homeostasis All living things, from single cells to entire organisms, have mecha- nisms that allow them to maintain stable internal conditions. Without these mechanisms, organisms can die. For example, a cell’s water content is closely controlled by the taking in or releas- ing of water. A cell that takes in too much water will rupture and die. A cell that doesn’t get enough water will also shrivel and die. Homeostasis (HOH-mee-OH-STAY-sis) is the maintenance of a stable level of internal conditions even though environmental conditions are constantly changing. Organisms have regulatory systems that maintain internal conditions, such as temperature, water content, and uptake of nutrients by the cell. In fact, multi- cellular organisms usually have more than one way of maintain- ing important aspects of their internal environment. For example, an owl’s temperature is maintained at about 40°C (104°F). To keep a constant temperature, an owl’s cells burn fuel to produce body heat. In addition, an owl’s feathers can fluff up in cold weather. In this way, they trap an insulating layer of air next to the bird’s body to maintain its body temperature. Metabolism Living organisms use energy to power all the life processes, such as repair, movement, and growth. This energy use depends on metabolism (muh-TAB-uh-LIZ-uhm). Metabolism is the sum of all the chemical reactions that take in and transform energy and materials from the environment. For example, plants, algae, and some bacteria use the sun’s energy to generate sugar molecules during a process called photosynthesis. Some organisms depend on obtaining food energy from other organisms. For instance, an owl’s metabolism allows the owl to extract and modify the chemi- cals trapped in its nightly prey and use them as energy to fuel activities and growth. Growth and Development All living things grow and increase in size. Some nonliving things, such as crystals or icicles, grow by accumulating more of the same material of which they are made. In contrast, the growth of living things results from the division and enlargement of cells. Cell division is the formation of two new cells from an existing cell, as shown in Figure 1-3. In unicellular organisms, the primary change that occurs following cell division is cell enlargement. In multi- cellular life, however, organisms mature through cell division, cell enlargement, and development. Development is the process by which an organism becomes a mature adult. Development involves cell division and cell differen- tiation, or specialization. As a result of development, an adult organism is composed of many cells specialized for different func- tions, such as carrying oxygen in the blood or hearing. In fact, the human body is composed of trillions of specialized cells, all of which originated from a single cell, the fertilized egg. This unicellular organism, Escherichia coli, inhabits the human intestines. E. coli reproduces by means of cell division, during which the original cell splits into two identical offspring cells. FIGURE 1-3 Observing Homeostasis Materials 500 mL beakers (3), wax pen, tap water, thermometer, ice, hot water, goldfish, small dip net, watch or clock with a second hand Procedure 1. Use a wax pen to label three 500 mL beakers as follows: 27°C (80°F), 20°C (68°F), 10°C (50°F). Put 250 mL of tap water in each beaker. Use hot water or ice to adjust the tem- perature of the water in each beaker to match the temperature on the label. 2. Put the goldfish in the beaker of 27°C water. Record the number of times the gills move in 1 minute. 3. Move the goldfish to the beaker of 20°C water. Repeat observations. Move the goldfish to the beaker of 10°C. Repeat observations. Analysis What happens to the rate at which gills move when the temp- erature changes? Why? How do gills help fish maintain homeostasis? Quick Lab mb06se_bios01.qxd 5/18/07 10:37 AM Page 8 THE SCIENCE OF LIFE 9 Reproduction All organisms produce new organisms like themselves in a process called reproduction. Reproduction, unlike other characteristics, is not essential to the survival of an individual organism. However, because no organism lives forever, reproduction is essential for the continuation of a species. Glass frogs, as shown in Figure 1-4, lay many eggs in their lifetime. However, only a few of the frogs’ off- spring reach adulthood and successfully reproduce. During reproduction, organisms transmit hereditary informa- tion to their offspring. Hereditary information is encoded in a large molecule called deoxyribonucleic acid, or DNA. A short segment of DNA that contains the instructions for a single trait of an organism is called a gene. DNA is like a large library. It contains all the books—genes—that the cell will ever need for making all the struc- tures and chemicals necessary for life. Hereditary information is transferred to offspring during two kinds of reproduction. In sexual reproduction, hereditary information recombines from two organisms of the same species. The resulting offspring are similar but not identical to their parents. For example, a male frog’s sperm can fertilize a female’s egg and form a single fer- tilized egg cell. The fertilized egg then develops into a new frog. In asexual reproduction, hereditary information from different organisms is not combined; thus the original organism and the new organism are genetically the same. A bacterium, for example, reproduces asexually when it splits into two identical cells. Change Through Time Although individual organisms experience many changes during their lifetime, their basic genetic characteristics do not change. However, populations of living organisms evolve or change through time. The ability of populations of organisms to change over time is important for survival in a changing world. This factor is also impor- tant in explaining the diversity of life-forms we see on Earth today.

Gereedschappen en materialen voor draden trekken - Starter Quiz

Related quizzes

Related quizzes